One of the difficulties in finding Chl d in nature was its overlap with Chl b on a reversed-phase HPLC trace. marina is composed of Chl d', pheophytin in PS II is not d-type but a-type ( Akiyama et al. In 1996, a unique cyanobacterium, Acaryochloris marina, with Chl d as the dominant chlorophyll was discovered in colonial ascidians ( Miyashita et al. 1), functions as the primary electron acceptor in PS II ( Klimov et al. 2001), and a metal-free Chl a, pheophytin (Phe) a ( Fig. 1), constitutes the primary electron donor of P700 in PS I as a heterodimer of Chl a/a' ( Kobayashi et al. Excitation of the primary donor reduces the primary and secondary electron acceptors, which again are often specially-tailored chlorophylls e.g., the 13 2-epimer of Chl a, Chl a' ( Fig. However, primary charge separation is initiated by a few specially-tailored chlorophylls in the reaction centers (RCs). 1) is the major chlorophyll in cyanobacteria, algae and terrestrial plants. Graduate School of Human and Environmental Studies, Kyoto University, Kyoto, JapanĬhlorophylls are essential components in oxygenic photosynthesis, and chlorophyll (Chl) a ( Fig.National Food Research Institute, Tsukuba, Japan.Research Center for Math and Science Education, Organization for Advanced Education, Tokyo University of Science, Tokyo, Japan.Institute of Industrial Science, University of Tokyo, Tokyo, Japan.Institute of Materials Science, University of Tsukuba, Tsukuba, Japan.
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